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Hylobates
About Gibbons,
Nomascus,
Symphalangus,
Hoolock
GENUS HYLOBATES
(Illiger, 1811)
The remaining gibbon species are classified in the genus Hylobates.
This genus has a diploid chromosome number of 44
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photo
by S. O'Reilly
Hylobates a. agilis (left)
& N. gabriellae x N. leucogenys siki
(hybrid)
(right) juvenile females--behavioral enrichment: gibbons playing with a
hat.
photo by A. Mootnick

H. albibarbis adult male
photo by A. Mootnick

H. agilis unko adult male
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Hylobates agilis (Cuvier, 1821). Agile gibbon.
General Description: H. agilis is commonly called the agile, dark-handed,
slender (a common name in the 1800s), or active gibbon. This species is
subdivided into three subspecies. Males and immature females have
whitish brows and cheek patches. The hair of the male's genital tuft is
50 mm long and generally the same color as, or slightly paler than, the
body hair (Marshall & Sugardjito, 1986). Females begin to lose their
cheek patches at six years of age. Adult females who have been housed in
low light conditions or who have nutritional deficiencies may lack the
white brow. The great call of the female agile gibbon is somewhat
similar to that of the female lar gibbon.
Hylobates agilis unko, the lowland agile gibbon, is highly threatened
and indigenous to eastern
Sumatra and the northern
peninsula of Malaysia.
Hylobates albibarbis, bearded
gibbon. Area: Southwestern Borneo.
The Hylobates albibarbis is light brown with dark-brown to brown-black
underparts, hands, feet, and cap and a white brow. H. albibarbis has
a song that has longer and slower notes when compared with Hylobates agilis
unko and H. a. agilis. Hylobates agilis unko and H. a. agilis are difficult
to distinguish from each other, while H. albibarbis is sometimes mistaken for
the northern Mueller's gibbon.
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photo by D.
Johanson

Adult female Hylobates a. agilis (mountain
agile gibbon) going through her final color change, with a 6 day old male
infant.
photo by A. Mootnick

H.
agilis agilis juvenile female
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photo by Lion Country Safari

H. klossii
female |
Hylobates klossii
(Kloss, 1929). Kloss' gibbon.
Area: Mentawai Islands.
General Description: Both sexes have black hair and remain this
color
at all life stages. Kloss' gibbons have short hair, a broad chest, and
long legs, thumbs, and great toes. The hair forming the genital tuft is
short. There is inter-digital webbing (Groves, 1972). Adults have a
small muzzle, and hair on top of the head is flat. Infants crown hair
stands erect. The song of the female Kloss' gibbon, particularly the
great call, is more similar to those of female pileated (H. pileatus)
and Mueller's (H. muelleri) gibbons than it is to female lar (H.
lar)
and agile gibbons (H. agilis). Kloss' gibbon populations are all
classified into one species, but there are some variations in hair
length, the direction of hair grown on the outer side of the forearm,
and body size on different islands (Groves, 1972, 1984).
Kloss' gibbon was originally described as a dwarf siamang (Miller,
1903). The confusion stemmed from the fact that all ages and both sexes
are black and the degree of webbing between the toes. Kloss (1929)
classified this species as an intermediate between the genera
Symphalangus and Hylobates. Schultz (1933), Groves (1972), and Tenaza
(1975) subsequently showed this species to be more closely related to
the species of the genus Hylobates, justifying their inclusion of Kloss'
gibbon within this genus. |
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photo by A. Mootnick

H. lar adult female |
Hylobates lar,
white-handed gibbon, are native to the forests of Myanmar, Thailand,
Yunnan, Malaysia and Sumatra. There are no established captive
breeding programs for this species, because there are approximately
300,000 in their native habitat, but they were recently found to be
extinct in China. |
photo by A. Mootnick

H. lar adult female & infant |
photo by S.
Kokel
H. moloch adult male
photo by S. Kokel

H. moloch adult male
photo by R.
Vera

H. moloch adult female
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Hylobates moloch (Audebert, 1798). Moloch, silvery, or Javan gibbon
General Description: Both sexes are silvery gray and have long, dense
hair at the neck, sides of the head, upper arms, and on the shoulders.
Either sex may have a distinct light-to-dark gray cap with the same
coloration occasionally under the brow (B3349). There is dark-gray hair
in the genital region. The adult females chest is sometimes partly
charcoal colored (Geissmann, 1995). Both sexes have white to white-gray
hair on the brow and surrounding the chin; this hair usually connects to
darker hair on the side of the face to form a face ring. Hair under the
chin grows upward, giving the appearance of a "goatee". Infants
are
lighter in color than are adults and change to silvery gray shortly
after birth (Groves, 1972). H. moloch is sometimes confused with
Mueller's gibbon because of similarities in coat color.
Hylobates moloch, Javan
gibbons, live in the fragmented forests of western and central Java, where
less than 2000 survive in the wild. GCC houses the only 11 Javan
gibbons in the USA, including the only Javan
gibbons ever born in the USA, and the only breeding pairs of Javan
gibbons in the Americas. Currently there are only five
institutions in the world that house breeding pairs of Javan gibbons.
CNN
article on
Javan gibbons
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photo by S.
Kokel

H. moloch adult male
photo by J. Zuckerman
H. moloch adult female
photo by S Kokel

H. moloch adult male
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photo by R. Wirth

H.
muelleri abbotti adult female & juvenile |
Hylobates muelleri (Martin, 1841). Mueller's, Bornean, or gray gibbon.
General Description: There are three subspecies. Adult male and
female
are identical in coat color, which varies from grey to grey-brown. The
hair of the adult male's genital tuft is 25 mm long (Marshall &
Sugardjito, 1986) and is typically darker than the body hair. Infants'
coats are lighter than their parents'. This species lacks a uniform
appearance in areas of geographic overlap with other Mueller's gibbon
subspecies. The great call of the female Mueller's gibbon is somewhat
similar to that of female pileated gibbons.
Eastern Mueller's gibbon (H. m. muelleri) and Abbott's gray gibbon
(H.
m. abbotti) are sometimes difficult to distinguish from moloch gibbons
because of similarities in coat color. H. m. funereus is occasionally
misidentified as the Bornean agile gibbon (H. agilis albibarbis) for the
same reason.
Hylobates muelleri abbotti (Kloss, 1929). Abbott's gray gibbon.
Area: Western Borneo. Abbott's gray gibbon has short, mouse-gray body hair with dark hair
in
the genital region. The brow is slightly paler than the head hair. Those
populations bordering H. muelleri funereus populations sometimes have
black hair on the throat and/or a slightly darker cap and upper chest
and inner aspects of the limbs, reflecting subspecific hybridization in
overlap zones.
Hylobates muelleri muelleri (Martin, 1841). Eastern
Mueller's gibbon. Area: Southeastern Borneo. The eastern Mueller's gibbon is pale grey with a
black cap, ventrum, hands, feet, and inner aspects of the limbs. It has a
thick white brow.
Hylobates muelleri funereus (Marshall & Sugardjito, 1986).
Northern Mueller's gibbon. Area: Northern Borneo. Northern Mueller's gibbon is dark gray or
gray-brown with black to blackish-brown cap, ventrum, throat, inner
aspects of the limbs, and genitals. Outer area of the lower limbs and tips
of the fingers and toes are generally paler or buffish-gray. Toes and
fingers are black on specimens from southern regions where the subspecies
overlap geographically with H. m. muelleri. (We will have a photo
in the near future.)
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photo by C. Whalley

H. pileatus
infant female
photo by
T. Motoyama

H. pileatus subadult male
photo by A.
Mootnick

H. pileatus adult male (left) & adult female (right) |
Hylobates
pileatus
(Gray, 1861). Pileated gibbon or capped gibbon.
General Description: This species is not divided into subspecies.
Adult
males have short, black hair with a thick, white brow band that becomes
thinner as it encircles the face. Males' fingers and toes are white,
with a slight fringe running halfway up the sides of the hands and feet.
There is a white prepubital patch. The crown cap is encircled by a
grizzled white streak on the sides of the head that becomes faint on the
back of the head. Some adult males have a faint gray grizzling on the
lumbar region. Adult females are silver-buff with a black heavily-furred
throat, inverted triangle on the ventrum that branches off to the
underarm area but stops short of the genital area. This black coloration
extends upwards to the bottom of the ears and narrows in front of the
ear to connect with the cap. The female's cap is large and black, with
long, silver-buff hair curved over the temples. Females have a thin,
white brow which sometimes extends laterally around the orbital ridge,
and there can be a trace of a white facial ring. Both sexes have a
lateral tuft along the crown's sides. Infants are slightly paler
silver-buff than are adult females. H. pileatus is the only species in
the genus Hylobates in which males undergo a color change (buff to
black). A male completed this transition in 6.5 years. males can begin
their color change at 18 months of age with a small black patch above
the sternum. The song of the female pileated gibbon is somewhat similar
to that of adult female Mueller's gibbon.
Hylobates pileatus,
pileated gibbons, are found only in SE Thailand (approximately 10,000), W.
Cambodia (approximately 10,000), and Laos (approximately 10,000). Half of
the pileated gibbon population in the Americas lives at GCC,
including the only two breeding pairs in the Americas. Currently there are only
twenty captive breeding pairs of pileated gibbons in the world.. "Pileated" means "capped".
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photo by A. Mootnick

H. pileatus adult female & infant female
photo by A. Mootnick

H. pileatus adult male |
About Gibbons,
Nomascus, Symphalangus,
Hoolock
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